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The insect collection of the Museum of Comparative Zoology is among the richest and historically most significant in North America, containing more than seven million specimens and the primary types of more than 33,000 species. The representation is worldwide and particularly strong in the major orders Coleoptera, Hymenoptera, Lepidoptera, and Diptera. Several historically important collections are housed at Harvard, including the type-rich beetle collections of Horn, LeConte, Melsheimer, Bowditch, Fall, and Darlington. The collections in many smaller orders are also among the most important in North America, i.e., Trichoptera, Odonata, Psocoptera, Neuroptera, and Collembola. The ant collection, alone comprising nearly a million specimens, is the largest and most important in the world. The fossil insect collection is the second most important one in the world. The MCZ Entomological collection was started in 1859 by Samuel Scudder and A.S. Packard, but includes the earlier material of T.W. Harris (the oldest intact North American synoptic insect collection), which was studied and partly determined by the pioneering American entomologist Thomas Say.